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From PewDiePie's Tuber Simulator Wiki
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Brains is one of the currencies in PewDiePie's Tuber Simulator. It helps upgrade items for the Knowledge Tree. Brains can be received by completing Quests, leveling up a theme or by leveling up your character. Brains are the only currency that you cannot get from Sponsor Eagle.

This button is shown on the Main Menu. By clicking the Knowledge Tree button, it will redirect you to the Knowledge Tree where you can spend your Brains.